Map Conception et réalisation : Jacques LE RENARD, B.I.M.M., M.N.H.N. Paris

CLEMAM
Check List of European Marine Mollusca 

Welcome to CLEMAM, an authoritative database of  the Marine Mollusca of Europe and surrounding areas


HELP
Scope
Editorial policy
Future developments
About CLEMAM
Clemam news
Interactive
Other links

Scope

     CLEMAM is a taxonomically oriented database of the marine Mollusca of Europe and adjacent areas (view the map of coastal sectors), aiming to a comprehensive coverage of the species in the eastern Atlantic from 26°N to the North Pole, the Mid Atlantic Ridge and the Mediterranean and Baltic Seas.
    The database lists the species currently accepted as valid (V, and boldface display), thence for each one its basionym (*, original reference where the specific name was introduced) and all synonyms (=) and questionable synonyms (?) known to the CLEMAM editor. Subsequent references to the species are not exhaustive; these include "chresonyms" (X, references to the species, using the valid name, or one of its synonyms or combinations), and misidentifications (!, references in which the valid name of another species is misapplied).
    Page references are given for all entries registered after 1995, and are being completed gradually for the initial batch, where only a statement of the name, author and date of publication were given.
    There are currently about 17.000 references, of which ca. 3500 are valid names.
    CLEMAM aims as being the standing reference for the systematics of European Mollusca, as well as a tool for species-oriented bibliographic search. The list of valid names of Mollusca in CLEMAM was contributed in 1999 to the European Register of Marine Species, an E.C. funded project led by Mark Costello (see the ERMS homepage), and we expect the CLEMAM framework to be the taxonomic base for other published and Internet checklists, catalogues and identification guides.

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Editorial policy
     The original contents of CLEMAM are cumulating those of four checkists published in paper format (see about CLEMAM). Nevertheless, this is not yet a complete coverage of all available names and the filling in of new material goes both to recording the current literature and to filling the backlog since 1758.
    Current literature, including all malacological journals and all publications known to the editors, is scanned for references of interest. Newly incorporated references have indication of authorship, relevant pages, and a comment on what the reference is about (in the style "original description", "reproductive biology"...). We include:
- all references which contain nomenclatural acts (descriptions of new species, designation of types).
- a selection of other references on the species present in CLEMAM area. Papers monographing anatomical or biological aspects of a species, and those where type material is illustrated, are usually listed, but choice may depend on how much attention the species receives (e.g. only major monographs will be listed for Mytilus) and on an editorial choice regarding the papers.
    As a rule, we try to keep CLEMAM as a recording tool and not as a place for reviewing or for introducing new data or new combinations of names. Thus, when a new taxon is published, it is deemed valid and leaved in its published combination and family placement until some conflicting view has been published. However, in some cases where published data are obviously irrelevant or wrong and would be misleading, a move is taken by the recorder; then a comment in square brackets is given to state which change has been made. This editorial filter is applied e.g. to records of exotic species from localities inside the CLEMAM area, which obviously are based on material transported by fishing boats and not on actually introduced populations.
    For some references where a decision on validity of a name would need revisional work out of the scope of the checklist, there are "incertae sedis" catchall references. Names listed under that heading are those that are nomenclaturally available, have no current usage, but must be considered in any taxonomic work concerned with the group.
    Coverage of the backlog is not systematic, due to the limited staff time available. Priority is given to gaps pointed by users, and to references containing nomenclatural acts.
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Some questions that you can ask CLEMAM:

What are the species of Nassariidae in the European area?
 
     Searching by higher taxa will yield a list of all valid species in the area belonging to this taxa, and thence their synonyms and chresonyms, but will not include records which are deemed incorrect (e.g. searching under Nassariidae will not yield Nassarius wolffi Knudsen, 1952, a tropical West African species which has two synonyms Nassa sadurnii and Hinia fringens malacitana, described from wrong localities in Spain; but those names are in CLEMAM and will be retrieved under the specific epithet).

What is the current valid name of Rissoa inconspicua?
Who is the author, and which is the date of publication of σσ?
 
    Searches under epithets or combinations are directed to questions such as "what is σσ ? ". If the request hits a synonym or otherwise invalid name, click on the name to go up to the full display and get the valid name, with its synonymy (including the searched name). Such searches will retrieve also the species which are erroneously cited in the area (with a comment stating that we reject the record).

    Searches under a binomen will retrieve only the exact binomen.
Warning: at this stage, CLEMAM does not contain many Genus/species combinations which have been widely used in the literature, but are neither the original binomen nor the current name (see "future developments"; e.g. we have Buccinum lapillus and Nucella lapillus, but not Purpura lapillus which has been used for over a century). Thus, it is better to search first the specific name (epithet); this will retrieve a list of the combinations in which it appears, including the valid one.
    Some names (e.g. forskoehlii, forskali; ruppelli, ruppellii, ruepelli) have elusive orthographic variants. Each variant is a totally different name in the base, so when in doubt, try them all or search under the appropriate genus or family name. If a species is registered as forskoehlii, search under forskoeli will yield "sorry, this taxon forsk... is unknown".

Where was this species published?
 
 For all entries posterior to 1995 and for many of the older ones, the author name in an entry is sensitive and gives access to the complete book or paper reference. These can be retrieved only one by one, but the user can copy and paste them in a wordprocessor document for his files.
 
 Now, you can also get bibliographic information directly, by typing the name (or the beginning of the name) of an author, instead of the name of a taxon. For instance, the "taxon or author name" Jeffreys will fetch the list of all the publications by Jeffreys, with, for each publication, a button pointing to the list of the taxonomic references present in that paper and which have been included in CLEMAM (primary references are in bold characters). Note : You may find difficulties with accents in names ; in many instances, this may be avoided by truncating the name (e.g., for Warén, type War) ...

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Future developments
     The main pending development of CLEMAM is the integration of geographical data. The area of coverage has been divided in sectors (view the maps of coastal sectors or of deep-water sectors), which are a compromise between political and biogeographical boundaries. So far, papers including only a citation of a species in a faunal list were not included in the database, but such publications will be the basis of validating a sector as part of the range of a species.
     There are short term plans to include references to as many as possible binomial combinations. We are aware that many combinations, which are neither the original binomen nor the currently accepted valid binomen, have been widely used in identification guides or in works on marine ecology. A link from these names to the valid names is crucial for the non-systematists who care for accuracy. Thus, we plan to include as chresonyms all combinations used in influent identification guides (e.g. Synopses of the British Fauna...) and names used as eponyms of biological communities.
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About CLEMAM
    The need to compile a check-list of the European marine molluscs was discussed during the 10th International Malacological Congress (Tübingen, 1989) and a resolution was passed to this effect by the General Assembly of Unitas Malacologica, and Anders Warén (Stockholm) and Philippe Bouchet (Paris) were asked to investigate into its feasability. The acronym CLEMAM (Check List of the European MArine Molluscs) was chosen because of its similarity to CLOFNAM and CLOFETA, the very successful regional checklists of Fishes compiled under the umbrella of UNESCO. In 1994, an ad hoc meeting, funded by Unitas Malacologica, brought together in Paris malacologists representing several regional or national lists of marine molluscs:

Clemam Group in Paris

(i) Riccardo Giannuzzi-Savelli, representing the Catalogo annotato dei molluschi marini del Mediterraneo, compiled by B. Sabelli, R. Giannuzzi-Savelli and D. Bedulli (1992) for Società Italiana di Malacologia, and published by Libreria Naturalistica Bolognese, Bologna.
(ii) Elizabeth Platts, representing the Checklist of British marine Mollusca, compiled by D. Heppell and S. Smith (1991), and published by the National Museums of Scotland.
(iii) Jon-Arne Sneli, representing the Annotated check-list of marine molluscs of the Norwegian coast and adjacent waters, compiled by T. Hoisaeter (1986) and published in Sarsia, 71: 73-145.
 In the following year, electronic versions of these three lists, in addition to a card catalogue of post-1960 names, were merged by Serge Gofas into a single list in text format. The resulting list was then transformed into database format by Jacques Le Renard, and the first CLEMAM was presented at the 12th International Malacological Congress (Vigo, 1995). This was completed by the data on Cephalopods of Angel Guerra (1992) published as Fauna Iberica vol. 1, and entered in 1996.

Who does what in CLEMAM?
Database design and development, web interface: Jacques Le Renard (Directeur de Recherches, INRA. E-mail: lerenard@mnhn.fr)
Taxonomic editor: Serge Gofas (Lecturer, University of Malaga. E-mail: sgofas@uma.es)
External liaison: Philippe Bouchet (Senior Curator, MNHN. E-mail: pbouchet@mnhn.fr).

Acknowledgements
We thank the following persons and institutions who have permitted the development of CLEMAM:
(1) Unitas Malacologica and its secretary (E. Gittenberger, Leiden, NL) and treasurer (J. van Goethem, Brussels, B) for seed money in the early phases of the project; Società Italiana di Malacologia and R. Giannuzzi-Savelli (Palermo, I), J.-A. Sneli and T. Hoisaeter (Trondheim and Bergen, N), E. Platts, D. Heppell and S. Smith (Edinburgh, UK), and A. Guerra (Vigo, SP) for making available electronic versions of published checklists.
(2) Maria-Antonietta and Paolo Angioy, editors of the magazine La Conchiglia (Roma), for permission to use pictures of Mediterranean gastropods published in the Atlante delle conchiglie marine del Mediterraneo.

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Clemam News
August, 1999:
    The great breakthrough for CLEMAM users this summer is that there is now access to complete bibliographic references, not only to the page references given with
taxonomic entries. The author name is now a sensitive zone, click on it and you should get a message: Publication Pxxxxx is: (complete reference). Of course, if the reference is
not yet in the base (about 50% of them are there), you only get the author name.
    Outstanding new entries! There is now a species of Bathymodiolus in (rather: at the edge of) CLEMAM area.
    The difficult Cephalaspid genus Diaphana has been revised
by Tom Schiøtte, so that some order has been brought.
    New direct links have been made to the test sheets of the forthcoming Atlas of Alien and Introduced species which is being prepared at the initiative of CIESM, Monaco.
Try references to Cerithium scabridum and to Anadara demiri (click on the words CIESM atlas in chresonym list; from there you can go to the CIESM home page ane read more about the atlas).
 
August, 2001:
Having run more than 58,000 requests, CLEMAM gets a brand new homepage...
 
November, 2001:
Now, you can get bibliographic information directly, by typing the name (or the beginning of the name) of an author, instead of the name of a taxon. For instance, the "taxon or author name" Jeffreys will fetch the list of all the publications by Jeffreys, with, for each publication, a button pointing to the list of the taxonomic references present in that paper and which have been included in CLEMAM (primary references are in bold characters). Note : You may find difficulties with accents in names ; in many instances, this may be avoided by truncating the name ...
 
End of July, 2002:
A huge catastroph : the whole interface software has been lost at once, due to a hard disk failure on the M.N.H.N. server. No backup copy of the interface software can be found at all !!! This means that Jacques Le Renard has to write all the software again, from scratch [for time saving, he'll do that on his personal web site, hence the strange URL].
Fortunately, the database itself and its content are safe (it has been copied on a mirror site).
 
September, 2002:
The M.N.H.N. server is still down. But here comes the new CLEMAM!
A new brand software has been written by Jacques, with a completely new design. It allows to navigate among taxonomic and bibliographic data more easily, and offers additionnal features, such as a multilingual support, and the ability to work with lists of names instead of single names. Moreover, CLEMAM inherits some original features of the FOSSILS database [also re-written by Jacques during his summer "hollidays"]: a geographical gazeteer (still to be filled in by Serge for CLEMAM).
 
February, 2003: The underlying software of CLEMAM has been incorporated into TAXIS (i.e., TAXonomic Information System), a new generic tool. Although TAXIS is particularly aimed at managing all the systematical aspects of collection databases, it is also used now, in a somewhat limited way, as the engine that powers the CLEMAM database. This explains why some new concepts have appeared here and there in the new CLEMAM interface. For an explanation of new terms used, see the TAXIS help page.
 
April, 2003: A management html interface has been added to TAXIS. Now, Serge Gofas, our expert in taxonomy, can freely work on the database, from anywhere in the world. Allthough this interface can be seen by everybody (example), only those with the right password can modify or add information to the database.
 
End of July, 2003:

NEW : If you want to use printouts of CLEMAM, you can now download printable systematic lists with PDF format. A new radio button has appeared on the SYSTEMATICS form, called 'pdf list'. When you select it and give the name of a taxon (of any rank), you'll get an Acrobat readable document, that can be printed easily and neatly. If the taxon you have entered is above the Order rank, the list does not include the species taxa; otherwise, the list goes down to the species level. [Due to time limitations imposed by the used server, it may happen that some lenghty lists are truncated before they have completed, resulting in fragmentary and thus unreadable files; this is particularly true when the server is overloaded.]
Ex.: List of MURICOIDEA

 
April, 2005: The server now runs a new php-5 platform, with improved performances.
 

Interactivity
     We welcome comments, suggestions of items to be included, and suggestions for corrections. Comments regarding the database structure should be addressed to Jacques Le Renard, comments regarding taxonomy should be addressed to Serge Gofas.

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Other links
    Some of te references in CLEMAM are direct species-level links to other internet sites of interest where the species is addressed (try links to Bernard Picton's Nudibranch site with many species of Doto, Flabellina...). However we keep those links down to a few dependable sites, so as to avoid too many brocken links with "error 404: file not found" messages.

    More general links to sites of interest can be found below

Nudibranchs of the British Isles.

MALACOLOG-3.0: A database of the Recent Gastropods of the Western Atlantic, based in Philadelphia and maintained by Gary Rosenberg.

MOLLIA: Information for malacologists, on the Berkeley website. Don't miss either the virtual exhibits of the Museum of Paleontology!

Mollusk research collections: A listing (but not a searchable database) of major malacological collections, by geographical area, by K. S. Cummings (Champaign).

Other connections related to Conchology (Guido Poppe)

Picture database of shells, by G. Poppe & Goto

Il Mediterraneo e le sue conchiglie: Shells from an aesthete's point of view, with some of the best quality photography on the web.

Catalogo interattivo: Molluschi del Pliocene Italiano.

Mediterranean slug site.
 

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